1Department of Cellular and Molecular Pharmacology, University of California, San Francisco, San Francisco, CA, USA.

2Howard Hughes Medical Institute, University of California, San Francisco, San Francisco, CA, USA.

4Biological and Medical Informatics Graduate Program, University of California, San Francisco, San Francisco, CA, USA.

5Integrative Program in Quantitative Biology, University of California, San Francisco, San Francisco, CA, USA.

6Center for Computational Biology, University of California, Berkeley, Berkeley, CA, USA.

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Single-cell lineages reveal the rates, routes, and drivers of metastasis in cancer xenografts - Science